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Complex animals: Annelids & arthropods

Hank continues our exploration of animal phyla with the more complexly organized annelida and arthropoda, and a biolography on insects. Created by EcoGeek.

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  • marcimus pink style avatar for user jiggly puff
    are there more complex animals then humans
    (10 votes)
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    • piceratops ultimate style avatar for user Just Keith
      I am not sure what you mean by "more complex" because that really isn't too useful a measure when comparing animals. There are certainly animals that have abilities we don't. For example, there are animals that can do such things as detect electromagnetic fields, sense north/south, inject venom, fly, see heat, use sonar, smell prey miles away, generate electric shock, etc.
      (25 votes)
  • blobby green style avatar for user sandeep dutt
    R/SIR, is moulting and ecdysis all the same?
    (9 votes)
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    • leafers seedling style avatar for user Darpan Raghav
      Yes, they are the actually the same. Moulting is kind of maturation or say transition from one stage of development to the other and this is brought in insects by hormones. One of the major hormone for bringing this change is Ecdysone. Thus the process is aka ecdysis. I hope its clear. :)
      (13 votes)
  • piceratops seed style avatar for user Sachin Kumbhar
    How does a dragonfly flap its wings?
    Can someone describe it as an mechanical movement pls?
    (7 votes)
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    • orange juice squid orange style avatar for user CarlBiologist
      There are little muscle fibers attached to either side of the wings. When the fibers are stimulated by the brain of the dragonfly they contract, pulling the wings in the direction (up or down) that the fibers are attached to. When the stimulation stops, the fibers relax. Then the brain stimulates the fibers on the other side of the wing to pull the wing in the opposite direction. The process repeats and the wing flaps up and down.
      (11 votes)
  • blobby green style avatar for user Nick Martain
    What actually evolutionarily pushed the human species to have 2 specialized genders instead of being simply hermaphrodites? Would it be because having 2 genital parts on each person would be very energy consuming? If so, why are there hermaphrodites at all?
    (2 votes)
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    • piceratops ultimate style avatar for user Just Keith
      Nature has produced several variations on sexual reproduction. It is an advanced topic that really you'd need an entire college level course to even begin to cover. However, there is a strong favoring of independent male and female sexes. This is so favored that it has apparently evolved independently more than once (in plants, in animals, and in some fungi).

      Just one of the issues thought to be driving this distinction is the organelles. When the organelles from different individuals are mixed into a zygote, they can prove to be incompatible. Thus, having the male gamete contain only the bare minimum needed and the female gamete contain everything else, tends to solve this problem.

      The separate sexes in animals arose hundreds of millions of years before humans evolved.
      (6 votes)
  • aqualine tree style avatar for user Revathinandaak
    OK, this may sound stupid but what is the actual difference between shrimps and prawns?
    (2 votes)
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    • old spice man green style avatar for user Matt B
      Haha, great question! :-) There is actually no difference:
      The terms shrimp and prawn have no definite reference to any known taxonomic groups. Although the term shrimp is sometimes applied to smaller species, while prawn is more often used for larger forms, there is no clear distinction between both terms and their usage is often confused or even reverse in different countries or regions. Prawn is a common name, used particularly in the United Kingdom, Ireland, and Commonwealth nations, for large swimming crustaceans or shrimp, especially those with commercial significance in the fishing industry.
      (5 votes)
  • female robot amelia style avatar for user Sarah Battista
    At around , he mentions that the wings are basically an extension of the cuticle of the thorax. Could you realistically say an arthropod is flying on its thorax's 'fingernails'?
    (3 votes)
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  • female robot ada style avatar for user 18bagaian
    Do oysters have tentacles?
    (3 votes)
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  • purple pi pink style avatar for user syd
    are molluscs any relations to this?
    (2 votes)
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  • aqualine seed style avatar for user G
    Is an earthworm viviparous?
    (2 votes)
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    • leafers ultimate style avatar for user Justin
      No. While earth worms reproduce sexually they do not produce young internally. Their young spawn from eggs that are fertilized externally on a structure known as the clitellum (the thick section visible on all earthworms). Earthworms are hermaphroditic although they cannot self fertilized their eggs so It is done by each organism execrating both sperm and egg from their proglottids and rubbing on each other. Once the eggs are fertilized they slide off the parent into the soil where they mature.
      (2 votes)
  • leaf green style avatar for user Jchen314
    How do barnacles have body segments?
    (2 votes)
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Video transcript

- Hi there, just hangin' out with some of my distant relatives here, and I gotta say, I've got a lot of respect for these guys, because they are some of the most successful organisms on the Earth. We think we run things on this planet, but we don't, they do. On one acre of cultivated land, annelids like this earthworm process about 16,000 pounds of soil a year, which makes plant life, and our lives, therefore possible. And not only are there about a billion, billion arthropods like this in the world, that's 10 to the 18th power, by the way, but also 80% of known animal species are arthropods. Now don't get me wrong, even though bugs and spiders and worms and shrimp totally outnumber us humans, we are far more complex than them when it comes to, like, arm-wrestling and guitar playing and long division we totally school them. But complexity like ours had to start somewhere. And it started with a very special trait that we share with these animals. Can you see the resemblance? (peppy rock music) Last time we talked about how in the very simplest animals there are two traits that indicate an animal's relative complexity. There's how many germ layer they develop when they're embryos and whether or not they have a coelom, or a body cavity that holds their organs. So it's in the next two phyla, Annelida and Arthropoda, where we find the new big thing in animal complexity: segmentation. Segmentation is the repetition of anatomically identical units that can be added to and modified to serve different purposes as animals evolve. And evolution is the way to win it, folks. In fact, the three biggest and most diverse groups of animals in the world are the ones that display segmentation: Annelida, Arthropoda, and Chordate, which includes the vertebrates. All segmented animals have a common ancestor that probably lived about 600 million years ago. That's how long ago it was when one of your grandparents and one of the earthworm's grandparents and one of the beetle's grandparents all played on the same softball team, pretty crazy. Segmentation has proven to be unbelievably useful from an evolutionary perspective. In humans, you see anatomically identical pieces repeated along an axis, from our butts to our heads. They can be a little hard to see because they're so highly modified, but think about our vertebrae, they're segments, our ribs are segments, the cartilage around our trachea, those are segments. Even the folds in our brains are segments. They're crazy evolved segments, but segments just the same. Among today's animal phyla, the earlier to display segmentation is Annelida, which includes leeches and earthworms and lugworms. See how their whole bodies look like rings fused together? Segments. In fact, the word Annelida comes from the Greek for little rings, and when you look at an annelid, you see that they're all really obviously segmented. Now this segmentation is a great example of synapomorphy in annelids. Synapomorphies are traits that set one group of animals apart from its ancestors and from other groups that came from the same ancestor. So unlike their flatworm and nematode cousins, annelids are segmented. And they've also got little bristles on their bodies, called chaetae, that provide traction and help them move through the dirt. These are both little extra somethings that annelids have that less complex relatives don't have, and that their common ancestors didn't have. Synapomorphies, literally shared, derived traits, are usually the defining traits of a phylum. But you can also learn something about an animal's lineage by comparing plesiomorphies: very basic traits that are shared by animals with a common ancestry. So between the Platyhelminthes and the nematodes and the annelids, one plesiomorphy is that they're all worm-shaped, which tells you that they have a common, distant ancestor that was wormy looking. So as we talk about these phyla and the classes within them, notice how they're similar and different from each other. For instance, within Annelida there are three different classes. Everybody's favorite, of course, is the Oligochaetes, the earthworms. Their name refers to the synapomorphy I just mentioned, they have chaetae, or bristles, but only an oligo, few of them. And they're everyone's favorite because they eat soil and crap it out the other end, allowing air and water to circulate in the soil. Plus, their poo is rich in things that plants need to grow, like nitrogen and calcium and magnesium and phosphorus, and now I'm gonna go watch my hands. Now on to the class Hirudinea, the leeches, a lot of which are parasitic and, you know, eat blood and other bodily fluids. The synapomorphy of leeches are their powerful suckers, they've got them on both ends of their bodies, the posterior one being used to anchor itself while the anterior one that surrounds its mouth attaches to its host. All leech species are carnivorous and they are hermaphrodites, like earthworms. Now, the Polychaetes are bristly worms, hence a synapomorphy of this class is their poly, many, chaetae, bristles. Almost all of these are marine species and they're really diverse, but the ones you've probably seen evidence of are lugworms, the ones that dig holes at the beach and leave piles of castings on the sand. Okay, I know, you've had enough of worms. And now we've got Arthropoda to talk about, and that's very easy to do because there are a lot of them. Like I said, they totally outnumber us. Just to put things into perspective, there are more insects in a square mile of rural land than there are human beings on the earth. One reason that scientists think arthropods do so well has to do with their, you guessed it, segmented bodies! Fossils of early arthropods showed that there used to be very little variation between the segments. But as they evolved, segments fused and became specialized for different functions, which led to crazy amounts of diversity. So much diversity that Arthropoda includes stuff like scorpions, butterflies, and lobsters, which I know, seems like maybe a bit of a stretch, but here are the synapomorphies that make them all arthropods. One, they have segmented bodes that, in most cases, are broken up into three segments: head, thorax and abdomen. And check this out, here segmentation in arthropods is a good example of plesiomorphy. It's a basic ancient trait that they share with annelids and us chordates, for that matter, dating back to that softball game our forebears played some 600 million years ago. Two, all arthropods have an exoskeleton, a hard outer shell made out of chitin, which is a really tough carbohydrate that's chemically similar to the cellulose that you find in plants. And, in order to grow bigger they actually have to shed it. And be glad that you don't have to do that, because it looks like kind of a traumatic experience for them. Finally, they've all got paired and jointed appendages, which is actually where their name comes from. Arthropod means jointed feet, but it's not just their legs that are jointed, some of them have claws and jointed antennae, and they all have these external mouthparts that are also jointed. So that's what arthropods have in common, but they are grouped into four subphyla based on how they differ from each other. First, and perhaps most terrifying, you have your Cheliceriformes, which include spiders and scorpions, but also horseshoe crabs, which are kinda nice, and ticks, which aren't, and mites, which don't bother you at all. Probably, they might. Cheliceriformes comes from the Greek for arm-lips, which, whatever Greeks, refer to their long, fang-like pincers. Unlike a lot of arthropods, they have simple eyes with just one lens and not compound eyes, like flies, and they lack antennae. Most Cheliceriformes are landlubbers, but the fossil record tell us that a lot of them were marine back in the olden days. Sea spiders and horseshoe crabs are the only ones left now. The largest class of Cheliceriformes are the arachnids, the group that includes spiders and scorpions and ticks and mites. And they have what's called a cephalothorax, which is a head segment and a thorax segment fused together with eight legs and an abdomen behind. Most arachnids are carnivorous or parasitic, and they're very skittery, just sayin'. Next, Myriapoda, or many feet, includes what you would expect, the millipedes and centipedes. These are where the arthropods were like, let's see how far we can take this segmentation thing, shall we? All myriapods are terrestrial and have antennae and sort of scary jaw-like mandibles. Millipedes are vegetarians, and they may have been some of the very first animals to live on land, where there were mosses and primitive vascular plants for them to munch. Also, although millipedes have a crap ton of legs, they don't have as many as a thousand, as their name implies. They've actually got anywhere between 94 and 394 legs, depending on the species. Centipedes, whose name is a little more apt, as they generally have between 20 and 350 legs, are carnivores and have poisoned claws to paralyze their prey. So if you're looking to cuddle with a myriapod I'm going to advise you to go with the millipede. But please, save some love for the Hexapoda, meaning six feet, most of which are insects. The synapomorphies they share are three part bodies consisting of a head, a thorax, and an abdomen, three pairs of jointed legs that come off the thorax, compound eyes, and two antennae. Now, think of any random way you could put these characteristics together, and you'd probably come up with something that exists, because you guys, there are so many damn insects out there, you have no idea. There are more species of insects than there are all other animal species combined. Which is why I'm taking this opportunity to do a Biolo-graphy, the insects edition. (cheery piano music) Very little is understood about the evolution of insects, but scientists think that they probably split off from their crustacean cousins about 410 million years ago. And for tens of millions of years, insects and some little skittery invertebrates were about the only land dwelling animals. About 320 million years ago, thanks to the high oxygen levels of the Carboniferous period, some insects grew to be terrifyingly big. Like the Meganeura, which looked like a dragonfly with a two-foot wingspan. But, since an insect's size is restrained by its respiratory system, as oxygen levels went down, these massive insects couldn't circulate enough oxygen to keep their gigantic bodies going, and they died off. The next major milestone of insect history occurred around 120 million years ago, which is when most flowering plants evolved. And with it, the sweet spirit of cooperation that insects and flowering plants still share to this day. In fact, some insects and flowering plants have co-evolved really neat pollination strategies so they basically evolved together. Which I think is really sweet. And as a result of insect pollination, flowering plants are now the overlords of the plant world. And thus, everything smells nicer and looks prettier. Thanks, insects. Insects are the only arthropods that have developed the ability to fly and it has served them well. Insects' wings are just extensions of the cuticle of the thorax, so unlike birds and bats, which have to sacrifice walking legs in order to evolve wings, insects are just as graceful on the ground as they are in the air. But in order to be so awesome, insects had to develop this crazy thing called metamorphosis. In partial metamorphosis, the young, called nymphs, look exactly like the adult of the species and undergo a series of molts which allows them to get bigger and bigger, but they look basically the same the whole time. Most of the types of arthropods do this and some insects, including grasshoppers and cockroaches. However, complete metamorphosis is a process unique to some insects that lets them completely change shape. Maggots turn into flies, mealworms turn into beetles, caterpillars turn into butterflies. The baby insect, called a larva, I have one right here, this is a rhinoceros beetle larva. (laughs) Pretty gross! Hangs out and eats until it's time to build a little case around itself, called a pupa. This is the rhinoceros beetle pupa, which is exceptionally creepy. (laughs) And then when it comes out of the pupa, it's fully grown rhinoceros beetle. It's like soft at first, and then its chitin hardens up and it becomes the adult. So insects, basically wizards, but not as delicious as Crustacea, the insects of the sea. Crustaceans include crabs, lobsters, shrimp, and barnacles, and like insects, they have three body segments, head, thorax, and abdomen. Although some have a cephalothorax, like spiders do. While most other arthropods have learned to love the land, very few crustaceans have. They have decided to put their energy into developing other amazing characteristics. For instance, lobsters and crayfish are like walking multi-tools, they have 19 pairs of appendages, some of which are claws, some are mouth parts, some are regular old walkin' legs. Some shrimp have evolved bioluminescence, which is pretty much the most amazing thing any animal can do as far as I can tell. And this yeti crab looks like a yeti! And we've covered most of the types of animals on the Earth in what, 10 minutes? Hopefully now you can see the resemblance between these guys. I love my caterpillar.