How CAM Plants are able to fix carbon at night so they don't have to keep their stomata open during the day. Created by Sal Khan.
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- How do CAM plants handle O2 without Bundle Sheath cells? How can they avoid photorespiration without Bundle Sheaths?(15 votes)
- It appears (from googling) that CAM plants DO photorespirate, and are NOT more efficient than C-3 plants.
Why CAM plants can't incorporate C-4 methods by just adding a Bundle Sheath is still a mystery to me.(15 votes)
- What is malate ? And what functions does it perform?(11 votes)
- Malate is a 4 carbon molecule that is produced during the Calvin-Benson cycle of a plant.(13 votes)
- i've heard a lot of people saying it's not good to place plants in bedrooms because "they release CO2 at night." the way i understand it is that plants, specifically ones doing C4 photosynthesis produce CO2 but release it to other cells within their structure and not in the air. are they wrong? or am i missing out on something?(6 votes)
- What happens is that a plant has mitochondira and chloroplasts, but the chloroplasts outnumber the mitochondria that even though the mitochondria produce carbon dioxide during the day during cellular respiration during the day, the hungry chloroplasts use all of it. But at night, the chloroplasts are not working so only the mitochondria are doing cellular respiration, giving you only co2.(12 votes)
- Large trees can produce many pounds of leaves over the year and drop them on the ground later. My question is, where does the material to make leaves come from. If is came directly from the soil, there would be a noticeable depression around the tree due to the large amount of material needed. Where does it come from?(4 votes)
- The vast majority of the material comes from the air. Carbon dioxide is extracted directly from the atmosphere, while atmospheric nitrogen is fixed in the soil by microorganisms. Water is extracted from the ground, and is replenished in a number of ways. Soil is mostly needed for some trace metals, ions, and phosphate.(12 votes)
- Sir, this video showed about fixation of CO2...but what about release of O2 in CAM plants how does it take place?(9 votes)
- after the malate is released in the vacuole in the cytosol its enters the chloroplast where NAD+ is reduced to NADH and yeild CO2 and pyruvate. CO2 is used in the calvin cycle, while pyruvate is convected to starch(0 votes)
- Why can't all plants do this? I mean if cams can store the malats then why can't the other plants?(3 votes)
- CAM plants are succulents and grow in dry regions. Due to excessive heat the stomata remains closed during the day to reduce transpiration. Their stomata opens at night. at night stomata are open by PEPA and OAA in the presence of PEPAcarboxylase. OAA gets reduced to malic acid and malate is accumalated during night. i.e. the acid concentration increases at night and decreases during day. In other plants the stomata opens during the day. And the other important factor is the RUBISCO which is activated in normal conditions which cannot be activated in CAMS due to high temperature and less carbon dioxide.(6 votes)
- At4:34Sal describes the way CAM plants fix carbon at night. But where does the energy to do this come from since there is no energy from photons? Is ATP saved up in the daytime for use at night? Is the cell burning glucose that was manufactured in the daytime?(4 votes)
- ATP is always present in the cell and being synthesized regardless of photosynthesis. Photosynthesis is not the only source of ATP and energy.(1 vote)
- so i dont understand how oxaloacetate acid is converted into malate?(2 votes)
- What are the other examples of CAM plants?
Are they helpful or harmful to us??(2 votes)
- CAM plants are commonly found in the desert, in hot and arid conditions. Some examples might be pineapple, or more commonly, cacti. I doubt you can classify them into Helpful or Harmful as a general category.(3 votes)
- Is it possible for plants to have photosynthesis at night.light reaction.....(2 votes)
A couple of videos ago, we saw that in classic C-3 photosynthesis-- and once again it's called C-3 because the first time that carbon dioxide is fixed, it's fixed into a 3-carbon molecule. But we saw the problem with C-3 photosynthesis is that the enzyme that does the carbon fixation, it can also react with oxygen. And when oxygen essentially reacts with ribulose biphosphate instead of your carbon, you get an unproductive reaction. Not only is it unproductive, it'll actually suck up your ATP and your NADPH and you'll go nowhere. So every now and then, when oxygen bonds here instead of a carbon dioxide, you get nothing produced, net. Everything becomes less efficient. And so in the last video, we saw that some plants have evolved a way to get around this. And what they do is, they fix their carbon on the outside, on cells that are actually exposed to the air. And then once they fix the carbon they actually fix it into a 4-carbon molecule, into oxaloacetate And then that gets turned into malate Then they pump the malate deeper within the leaf, where you aren't exposed to oxygen. And then they take the carbon dioxide off the malate, and this is where they actually perform the Calvin cycle. And even though you do have your rubiscos still there, your rubisco isn't going to have-- the photorespiration is not going to occur. Because it only has access to carbon dioxide. It does not have access to this oxygen out here. Now that's a very efficient way of producing sugars. And that's why some of the plants that we associate with being very strong sugar, or even ethanol producers, all perform C-4 photosynthesis. Corn, sugarcane, and crab grass. And these are all very, very efficient sugar producers. Because they don't have to worry too much about photorespiration. Now some plants have a slightly different problem. They're not so worried about the efficiency of the process. They're more worried about losing water. And you can imagine what plants these are. These are plants that are in the desert. Because these stomata, these pores that are on the leaves, they let in air, but they can also let out water. I mean, if I'm in the rainforest, I don't care about that. But if I'm in the middle of the desert, I don't want to let out water vapor through my stomata. So the ideal situation is, I would want my stomata closed during the daytime. This is what I want. So I want-- if I'm in the desert, let me make this clear. If I'm in the desert I want stomata closed during the day. For obvious reasons. I don't want all my water to vaporize out of these holes in my leaves. But at the same time, the problem is that photosynthesis can only occur during the daytime. And that includes the dark reactions. Remember, I've said multiple times, the dark reactions are badly named. They're more the light independent reactions. But they both occur simultaneously-- the light independent and light dependent-- and only during the daytime. And if your stomata is closed, you need to perform photosynthesis, especially the Calvin cycle, you need CO2. So how can you get around this? If I want to close my stomata during the day, but I need CO2 during the day, how can I solve this problem? And what desert plants, or many desert plants, have evolved to do, essentially does photosynthesis, but instead of fixing the carbon in outer cells and then pushing it in to inner cells and then performing the Calvin cycle, instead of outer and inner cells, they do it at the nighttime and in the daytime. So in CAM plants-- and these are called CAM plants because, I could tell you what it stands for. It stands for crassulacean acid metabolism. And that's because it was first observed in that species of plants, the crassulacean plant. But these are just called, you could call it CAM photosynthesis or CAM plants. They're essentially a subset of C-4 plants. But instead of performing C-4 photosynthesis, kind of an outside cells and inside cells, they do it at the nighttime and the day. And what they do is, at night they keep their stomata open. And they perform, and they're able to fix-- and everything occurs in the mesophyll cells and the CAM cells, in the CAM plants. So at nighttime, when they're not afraid of losing water-- let's say this is a mesophyll cell right here-- my stomata is open. Let's say that this is my stomata right there. And so it lets in carbon dioxide. I'm not worried about losing water vapor. It's night time right now. So carbon dioxide comes in here. And then it fixes the carbon dioxide. It fixes it the exact same way that the C-4 plants do. So you have your CO2 come in. You have your PEP . It's all facilitated by PEP carboxylase That's the enzyme. That can only fix CO2, that can only react with CO2, not with oxygen. And then that is used to produce-- and we saw it here in our CAM-4 diagram in the last video, that is to used to produce malate. A 4-carbon molecule. And then the malate-- and then this is what's key-- the malate get stored in other organelles in the cell. In the vacuoles , which are, you can kind of view them as large storage containers in the cell. So I drew this as the whole cell. I mean, this is actually all occurring in your chloroplast. But you can imagine your cell having a big storage center where the malate gets stored at night. And you can view malate as almost a carbon dioxide store. Because later on we can access the malate and get the carbon dioxide. And that's exactly what these CAM plants are going to do. So this is nighttime. Then the sun comes up. So now we're in the daytime. This desert plant, well maybe it's a cactus, it doesn't want to lose its water vapor. So it closes its stomata. This particular stoma now is closed. It's now closed. And you say, oh boy, how is it going to perform photosynthesis? Well, it can perform photosynthesis in that very same cell. Because it stored up all of this malate at night. And so now the malate can be pumped out of the vacuoles into the stroma of our chloroplast. And then you can have pyruvate break off. But the more important thing is you have CO2 break off. So you have a ready supply of CO2. And now we can perform our standard Calvin cycle. And in an environment only with CO2, our stomata is closed, so we're ready to go. Our CO2 reacts with ribulose bisphosphate It is catalyzed by rubisco It's the whole Calvin cycle and we produced our sugar. So this is kind of a neat adaptation. In these high, very efficient sugar-producing plants that aren't worried about water, they perform carbon fixation on things that are exposed to the air and then they pump kind of a stored version of the carbon deeper into the leaf to actually perform the Calvin cycle so that it's not lossy, so that photorespiration doesn't occur. Because down here you have no oxygen. The desert plants benefit from that property as well, but their whole concern is, I don't want to keep my stomata open in the daytime. So what I do is, I fix my carbon at night. But I use the exact same process. I use PEP carboxylase And I store my carbon dioxide at night. And in the daytime, I can actually-- when my light-dependant reactions are occurring, they're producing my ATP and my NADH I can also perform my dark reactions in the daytime. As I said, the dark reactions always occur in the daytime. Or my light-independent reactions. Because even though my stomata is closed, I have a store of carbon dioxide in the form of malate.